Urban Greylag Geese Anser anser nesting on roofs in the city of Hamburg, northern Germany

Main Article Content

Simon Hinrichs
Jean-Louis Berthoud
Friederike Woog

Abstract

Greylag Geese Anser anser usually nest on the ground, close to water. In Hamburg, northern Germany, a pair of urban Greylag Geese was first seen nesting on a roof terrace of a building, 25 m above ground level, in 2013. Since then, the number of breeding pairs showing this unusual behaviour has steadily increased to 21 pairs nesting on roofs in the city in 2024. Hatchlings appeared unable to leave leave these new nesting locations on their own and required human intervention to reach ground level. This type of nesting behaviour now seems to have become established in Hamburg’s Greylag Geese. As far as we know, there have been no previous reports of this behaviour in other towns or cities, and it therefore represents a new aspect in the species’ urban ecology.

Article Details

How to Cite
Hinrichs, S., Berthoud, J.-L., & Woog, F. (2025). Urban Greylag Geese Anser anser nesting on roofs in the city of Hamburg, northern Germany. Wildfowl, 75. Retrieved from https://tidsskrift.dk/Wildfowl/article/view/162737
Section
Standard Papers

References

Armstrong, T.E., Armstrong, D. & Armstrong, T. 2017. Observations on multiple-year tree-nesting by Canada Geese in southern Ontario. Ontario Birds 35: 105–114.

Askren, R.J. 2021. Movement ecology and habitat use of Canada geese using major metropolitan areas in the context of human-wildlife conflicts. Ph.D. thesis, University of Illinois, USA.

Auguie, B. 2017. gridExtra: Miscellaneous Functions for “Grid” Graphics R package version 2.3. R Foundation for Statistical Computing, Vienna, Austria. Available at https://CRAN. R-project.org/package=gridExtra (last accessed 26 September 2025).

Bates, D., Maechler, M., Bolker, B. & Walker, S. 2015. Fitting linear mixed-effects models using lme4. Journal of Statistical Software 67: 1–48.

Coath, M. 2006. Greylag Goose nesting in pine tree. British Birds 99: 365.

Dingemanse, N.J., Kazem, A.J., Réale, D. & Wright, J. 2010. Behavioural reaction norms: animal personality meets individual plasticity. Trends in Ecology and Evolution 25: 81–89.

Dinsmore, S.J., White, G.C. & Knopf, F.L. 2002. Advanced techniques for modeling avian nest survival. Ecology 83: 3476–3488.

Dorai-Raj, S. 2022. binom: Binomial Confidence Intervals for Several Parameterizations R package version 1.1-1.1. R Foundation for Statistical Computing, Vienna, Austria. Available at https://cran.r-project.org/web/packages/ binom/index.html (last accessed 26 September 2025).

Duncan, A., Duncan, R., Rae, R., Rebecca, G.W. & Stewart, B.J. 2001. Roof and ground nesting Eurasian Oystercatchers in Aberdeen. Scottish Birds 22: 1–8.

Fisk, E.J. 1978. The growing use of roofs by nesting birds. Bird Banding 49: 34–141.

Glutz von Blotzheim, U.N. 1990. Handbuch der Vögel Mitteleuropas. Band 2: Anseriformes. Teil 1. Aula Verlag, Wiesbaden, Germany. [In German.]

Groom, Q.J., Adriaens, T., Colsoulle, C., Delhez, P. & Van der Beeten, I. 2020. Site selection by geese in a suburban landscape. PeerJ 8: e9846.

Hart, L.A. & Downs, C.T. 2020. Greylag Goose (Anser anser Linnaeus, 1758). In C.T. Downs & L.A. Hart (eds.), Invasive Birds: Global Trends and Impacts. CAB International, Wallingford, UK.

Hartig, F. 2024. DHARMa: Residual Diagnostics for Hierarchical (Multi-Level/Mixed) Regression Models R package version 0.4.7. R Foundation for Statistical Computing, Vienna, Austria. Available at https://cran.r-project.org/web/ packages/DHARMa/ (last accessed 26 September 2025).

Hölzinger, J., Kroymann, B. & Kroymann, L. 2004. Graugans (Anser anser) brütet auf schilfgedecktem Haus am Max-Eyth-See in Stuttgart. Ornithologische Jahreshefte Baden-Württemberg 20: 101–107. [In German.]

Hoff, H.-J. 2005. Eine farbig beringte population von Graugänsen (Anser anser) in Öjendorf/ Hamburg. Hamburger Avifaunistische Beiträge 33: 157–173. [In German.]

Horák, P. 1999. Populace husy velké (Anser anser) hnízdící na stromech vyhynula. Crex – Zpravodaj Jihomoravské pobocˇky Cˇ SO 14: 41–45. [In Czech with English summary.]

Horal, D. & Heralt, P. 2012. Další prˇípad stromového hnízdeˇní husy velké (Anser anser). Crex – Zpravodaj Jihomoravské pobocˇky CˇSO 31: 155–157. [In Czech with English summary.]

Kleefstra, R. & Bles, A. 2000. Boomnesten van Grauwe Ganzen Anser anser: een nieuwe trend? Limosa 73: 148–150. [In Dutch with English summary.]

Kreutzkamp, I. 1996. Die Graugans (Anser anser) als brutvogel in Hamburg und umgebung. Hamburger Avifaunistische Beiträge 28: 129–158. [In German with English summary.]

Kreutzkamp, I. 2003. Die entwicklung der brutpopulationen von Graugans (Anser anser), Kanadagans (Branta canadensis) und Nilgans (Alopochen aegyptiacus) im Hamburger berichtsgebiet von 1990 bis 2002. Hamburger Avifaunistische Beiträge 32: 153–186. [In German with English summary.]

Lebeda, C.S. & Ratti, J.T. 1983. Reproductive biology of Vancouver Canada Geese on Admiralty Island, Alaska. The Journal of Wildlife Management 47: 297–306.

MacKey, D.L., Matthews, W.C. & Ball, I.J. 1988. Elevated nest structures for Canada Geese. Wildlife Society Bulletin 16: 362–367.

Mai, S., Berthoud, J.L., Haag, H. & Woog, F. 2022. Factors limiting reproductive success in urban Greylag Geese (Anser anser). PeerJ 10: e13685.

Mainwaring, M.C. 2015. The use of man-made structures as nesting sites by birds: A review of the costs and benefits. Journal for Nature Conservation 25: 17–22.

Mitchell, C., Black, J.M. & Ans, M.E. 1998. Breeding success of cliff-nesting and island-nesting barnacle geese in Svalbard. Norsk Polarinstitutt Skrifter 200: 141–146.

Nelson, U.C. 1953. Cliff-nesting Canada Geese on the arctic slope of Alaska. The Journal of Wildlife Management 17: 536–536.

Nieß, H. 1967. Graugänse in Hamburg. Vogel und Heimat 16: 134. [In German.]

Osiejuk, T.S. & Kuczyn´ski, L. 2007. Factors affecting flushing distance in incubating female greylag geese Anser anser. Wildlife Biology 13: 11–18.

Owen, M. 1980. Wild Geese of the World. Batsford, London, UK.

Posit Team. 2025. RStudio: Integrated Development Environment for R Posit Software, Boston, USA. Available at https://posit.co/products/ open-source/rstudio/ (last accessed 26 September 2025).

Robinson, D., Hayes A. & Couch S. 2025. broom: Convert Statistical Objects into Tidy Tibbles R package version 1.0.8. R Foundation for Statistical Computing, Vienna, Austria. Available at https://cran.r-project.org/web/ packages/broom/ (last accessed 26 September 2025).

Shaffer, T.L. 2004. A unified approach to analyzing nest success. The Auk 121: 526–540.

Shearer, D.J., Carter, T.C. & O’Neal, B.J. 2022. Canada geese (Branta canadensis) nesting on elevated structures in urban Indiana, USA. Ecology and Evolution 12: e8735.

Soldatini, C., AlboresBarajas, Y.V., Mainardi, D. & Monaghan, P. 2008. Roof nesting by gulls for better or worse? Italian Journal of Zoology 75: 295–303.

Wickham, H. 2007. Reshaping data with the reshape package. Journal of Statistical Software 21: 1–20. Available at http://www.jstatsoft.org/ v21/i12/ (last accessed 26 September 2025).

Wickham, H. 2011. The split-apply-combine strategy for data analysis. Journal of Statistical Software 40: 1–29. Available at https://www. jstatsoft.org/v40/i01/ (last accessed 26 September 2025).

Wickham, H. 2016. ggplot2: Elegant Graphics for Data Analysis Springer-Verlag, New York, USA. Available at https://ggplot2.tidyverse.org (last accessed 26 September 2025).

Wickham, H., François R., Henry L., Müller K. & Vaughan D. 2023. dplyr: A Grammar of Data Manipulation R package version 1.1.4. R Foundation for Statistical Computing, Vienna, Austria. Available at https://CRAN. R-project.org/package=dplyr (last accessed 26 September 2025).

Wickham, H., Vaughan D. & Girlich M. 2024. tidyr: Tidy Messy Data R package version 1.3.1. R Foundation for Statistical Computing, Vienna, Austria. Available at https://cran. r-project.org/web/packages/tidyr/ (last accessed 26 September 2025).

Woog, F. 2025. Behavioural ecology of urban Greylag Geese Anser anser – natural selection in the contemporary European city environment. Wildfowl 75: 3–31.